. Carnegie Institution of Washington publication. 84 THE WATERLILIES. The structure of the filament resembles that of the inner petals. The epidermis bears hair-bases, the parenchyma is made of soft, rounded cells, and air-spaces are present. There are one to three or more longitudinal vascular bundles. The mature anther is covered by a single layer of square cells, a little deeper than wide, with thickened inner and lateral walls ; the outer wall is unthickened. The anther cells are parallel and introrse, with longitudinal dehiscence (Fig. 37, a, b}. In Lotos and Brachyceras the division betw
. Carnegie Institution of Washington publication. 84 THE WATERLILIES. The structure of the filament resembles that of the inner petals. The epidermis bears hair-bases, the parenchyma is made of soft, rounded cells, and air-spaces are present. There are one to three or more longitudinal vascular bundles. The mature anther is covered by a single layer of square cells, a little deeper than wide, with thickened inner and lateral walls ; the outer wall is unthickened. The anther cells are parallel and introrse, with longitudinal dehiscence (Fig. 37, a, b}. In Lotos and Brachyceras the division between the two loculi of each cell is quite evident after dehiscence, as the pollen dust remaining after most has been shed lies in two pairs of grooves on the inner face of the anther. Chifflot (1902) has recently made a minute study of sporo- phyll structure in Nymphaea- ceae, including nine species of Nymphaea. He finds slight differences in the venation of the stamens. He sums up his work (which I have not had time to verify) in a classifica- tion based on the distribution of idioblasts (" sclerites ") and stomata in the stamen, and of idioblasts in the carpels (1. c., p. 280). We quote this in translation. The names are correct and mostly well chosen, unless in the case of "Ar. ; The material of this came from the Kew Herbarium, where N. rudgeana occurs under the name of TV. blanda, and is much more plentiful. SYNCARPIAE : Idioblasts rare in the filament, always present in the anther a. Idioblasts absent from filament, anther, and ovary (Xanthantha) A', flai'a. Idioblasts present in the filament and outer part of ovary (Castalia) c. a. Stomata present on upper epidermis of filament (Lotos). Idioblasts absent from mesophyll of ovary, present in carpellary styles A', thermalis. a. Stomata wholly absent from the stamen (Hydrocallis) b. b. Idioblasts of but one kind, present in filament and anther "N. ; b. Idioblasts of two kinds, local
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